Who explained the species-area relationship?

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(B) The species-area relationship was proposed by the German naturalist and geographer $Alexander$ $von$ $Humboldt$. He observed that within a region,

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Alexander von Humboldt

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(B) The species-area relationship was proposed by the German naturalist and geographer $Alexander$ $von$ $Humboldt$. He observed that within a region, species richness increased with increasing explored area, but only up to a limit. The relationship is described by the equation $S = CA^z$, where $S$ is species richness, $A$ is area, $Z$ is the slope of the line (regression coefficient), and $C$ is the $Y$-intercept.

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