Linkage was first studied by .....

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(C) Linkage was first observed and studied by $W. 	ext{ Bateson}$ and $R.C. 	ext{ Punnett}$ in the sweet pea plant $(Lathyrus 	ext{ odoratus})$.
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Bateson and Punnett

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(C) Linkage was first observed and studied by $W. 	ext{ Bateson}$ and $R.C. 	ext{ Punnett}$ in the sweet pea plant $(Lathyrus 	ext{ odoratus})$.
They observed that the genes for flower color and pollen shape did not show independent assortment as expected by Mendel's laws.
Although $T.H. 	ext{ Morgan}$ later provided the chromosomal theory of linkage and performed extensive work on $Drosophila$, the initial discovery and study of the phenomenon of linkage are credited to $Bateson$ and $Punnett$.

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